Education – University of Copenhagen

CCS > Education


Pre- and postgraduate courses

photo of students in the labThe CCS aims to provide a dynamic and friendly research environment for students at various levels both within our dedicated laboratories and on our university courses.

The CCS group leaders, postdocs and PhD students are participating in teaching on courses including: Molecular Biology, Medical Cell and Tissue Biology (Block D), Medical Genetics, Human Genetics, Molecular biology and Genetics , Advanced Live Cell Imaging, Reproductive Biology Course, and regular teaching seminars in genomics for embryologists (Region H).

Bachelor project, MSc and PhD defenses



  • 2nd Dec. 2019, Divya Achuthankutty will defend her PhD thesis: “Dissecting replication stress signaling: New regulatory mechanisms and experimental approaches” Main supervisor: Prof. Niels Mailand. Co- supervisor: Prof. Ian Hickson
  • 21st Nov. 2019, Andrés José Bueno Venegas will defend his PhD thesis: “Towards Defining the Essential Role of the SMC5/6 Complex in Human Cells”. Supervisor: Prof. Ian Hickson


  • 29th Aug. 2019, Karina Gaardahl passed her master thesis: “Investigating functional interactions between BRCA1 and RASAL2 in breast cancer cell lines”. Main Supervisor: Asso. Prof. Andres J Lopez-Contreras. Co-supervisor: Dr. Eliene Albers.
  • 30th Aug. 2019, Smaragdi Kompocholi passed her master thesis: “Further unravelling the role of ZGRF1”. Supervisor: Prof. Michael Lisby
  • 14th Aug. 2019, Keerthana Stine Viswalingam passed her maser thesis: “Characterization and Epistasis Analysis of a Novel Human Helicase ZGRF”. Supervisor: Prof. Michael Lisby
  • 4th Jul. 2019, Lorenza Garribba defended her PhD thesis: “Analysis of folate deficiency-induced genome instability in human cells”. Main Supervisor: Asso. Prof. Ying Liu. Co-supervisor: Prof. Ian Hickson.
  • 28th Jun. 2019,
    • Oline Jensen passed her bachelor project thesis: “The role of Mte1 in DNA repair”. Supervisor: Prof. Michael Lisby
    • Vanessa Dahl passed her bachelor project thesis: “Role of Sgs1/BLM protein acetylation in cellular ageing and DNA repair”. Supervisor: Prof. Michael Lisby
    • Simon Wenneberg passed his bachelor project thesis: “Development of an FN3 nanobody against the acetylated Sgs1 helicase”. Supervisor: Prof. Michael Lisby26th Jun. 2019, Ditte Marie Storm passed her bachelor project: "The effect of Topoisomerase II inhibition in chromosome stability”. Project supervisor: Prof. Eva Hoffmann.
  • 21st May 2019, Wei Wu defended his PhD thesis: “RTEL1 resolves conflicts between DNA replication and Transcription”. Main Supervisor: Asso. Prof. Ying Liu. Co-supervisor: Prof. Ian Hickson
  • 29th Mar. 2019, Eliene Albers defended her PhD thesis: “Generation and characterization of a PICH deficient mouse model.” Main Supervisor: Asso. Prof. Andres J Lopez-Contreras. Co-supervisor: Prof. Ian Hickson
  • 21st Mar. 2019, André Brannvoll defended his PhD thesis: “Guardians of the genome”. Main Supervisor: Prof. Michael Lisby. Co-supervisors: Asso. Profs Hocine W. Mankouri and Andrés López-Contreras.


  • 12th Dec. 2018,  Amruta Shrikhande defended her PhD thesis: "Understanding the regulation of endogenous mutagenesis”.(Brighton, University of Sussex) Main supervisor: Prof. Eva Hoffmann.  Co-supervisor: Dr Francis Pearl.
  • 20th Sept. 2018, Aiste Aleliunaite defended her PhD thesis: “Cellular response to site-specific replication blocks in mammalian cells”. Supervisor: Prof. Ian Hickson.
  • 25th Jun. 2018, Ireen Kooij passed her bachelor project: "Identification of Candidate Gene(s) Whose Mutation Predisposes to Radiation Sensitivity”.  Supervisor: Asso. Prof. Ying Liu. 
  • 10th May 2018,  Robert Blanshard defended his PhD thesis: "An investigation of genome stability in human female meiosis by genome wide chromosome fingerprinting and copy number analysis”. (Brighton, University of Sussex) Main supervisor: Prof. Eva Hoffmann.  Co-supervisor: Prof Tony Carr.
  • 17th Apr. 2018, Signe Winther Jørgensen defended her PhD thesis "Analysis of DNA replication stress at a telomeric region". Main Supervisor: Prof. Ian David Hickson. Co-supervisor: Asso. Prof. Hocine William Mankouri.


  • 29th Sept. 2017, Victoria Alexandra Bjerregaard defended her PhD thesis "Folate deficiency drives missegregation of the FRAXA locus". Main Supervisor: Asso. Prof. Ying Liu. Co-supervisor: Prof. Ian David Hickson.
  • 13th Sept. 2017, Özgun Özer defended her PhD thesis ”Telomere Instability in Human Cells - replication stress-induced mitotic DNA synthesis at difficult-to-replicate loci”.  Main Supervisor: Prof. Ian David Hickson, Co-supervisors: Asso. Profs. Ying Liu, Hocine William Mankouri and Claus Storgaard Sørensen


  • 31st May 2016, Ganesha Pitchai, defended his PhD thesis "Biophysical and Structural Characterization of the Interaction Between PICH and BEND3". Main Supervisor: Prof. Ian Hickson. Co-supervisors: Dr. Werner W. Streicher and Prof. Guillermo Montoya.


  • 28th Apr. 2015, Christian Friberg Nielsen, defended his PhD thesis "PICH Stimulates Topoisomerase IIa to Promote Sister Chromatid Disjunction in Mitosis” Main PhD supervisor: Prof. Ian Hickson. Co-supervisor: Asso. Prof. Ying Liu